Quantification of hierarchical substructure detected through seven major lines: intra-, inter-cluster allele sharing. In general, mean intra-cluster allele sharing increased over the course of the hierarchy and was always greater than the allele sharing observed among sibling clusters. The 95 per cent confidence intervals for mean allele sharing statistics are also shown and most often did not overlap for inter- and intra-cluster measurements. These data gave another indication of the high productivity of the partitions imposed by structure, even at fine levels of resolution. (Note: the margin of error is wide in the Kalash line, due to the small number of clusters (K = 3) defined for the sample.).